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In 1914, hoofer George Doane meets Eddie Martin at the Miner's Bowery Theatre when Eddie performs in an amateur contest. When the audience begins to heckle Eddie, George offers encouragement and stage directions from the wings. As a result of George's coaching, Eddie wins first prize, and George invites him to celebrate at Kelly's Café, a gathering place for entertainers. At the café, the womanizing George notices Constance Ford, who is sitting with her sister, Joan Mason, and their agent, Charlie Lucas. After introducing himself to Connie, George begins to dance with her to the tune of "It Had to Be You." They are interrupted by the appearance of Nancy Gaye, George's jealous girl friend, who insists that he leave with her. That night, George invites Eddie to join his burlesque act, and at the end of the season on the road, George decides to ditch Nancy in Maine while he goes to New York to look for Connie. In New York, George and Eddie return to the café, where they meet Connie, Joan and Charlie. When George asks Charlie about breaking into vaudeville, Charlie discourages him until Connie suggests combining her and Joan's sister act with George and Eddie's burlesque show. The new team of Doane, Martin, Mason and Ford is a success. On the road, Joan proposes to Eddie, who she says reminds her of a cocker spaniel, and George proposes to Connie, but their offers are rejected. In Baltimore, Nancy catches up to George, but Eddie ushers her out of the theater before she can see him. When Charlie criticizes their act as not being flashy enough to win a New York booking, George decides to raise the $10,000 needed to expand the show by breaking the team into five separate acts. After an exhausting season, they have saved $5,000, which George sends to Nancy when he learns that she has been involved in a serious accident and needs the money to recuperate. Afterward, Connie finally accepts George's proposal, and the team travels to Boston. Just as they raise the $10,000 needed to mount a production at the Palace Theatre in New York, Connie announces that she is pregnant. After their New York success, Connie takes a temporary leave of absence from the act while Joan, Eddie and George go back on the road. They are appearing on the same bill with Nancy when word comes that Connie has gone into labor. Joan and Eddie rush back to the city to be with Connie, leaving George to drive back with Nancy. Nancy, who is still in love with George, deliberately misses the New York turnoff, and as a result, George arrives at the hospital hours after Connie has delivered her baby and learns that the infant has died and Connie plans to divorce him. With the advent of World War I, Eddie goes overseas to entertain the troops and is reunited with George when he performs in a hospital where George is convalescing. At war's end, Charlie welcomes George home with a party at a nightclub. At the club, Connie tells George that she has accepted Charlie's proposal. Later, Connie begins to have doubts about the marriage and tells Charlie that she wants to return to work. As Connie takes to the road singing "It Had to Be You," George also hits the road, but his career takes a downward spiral and he is finally reduced to singing for drinks. Back in New York, when Charlie tells Eddie that Ziegfeld wants his and Joan's act, Eddie inquires about George. After Charlie replies that George has been spotted singing in a San Francisco speakeasy, Eddie, who is convinced that Connie still loves her ex-husband, decides to find his old friend. In San Francisco, Eddie locates George and pretends to be an alcoholic in need of his friend's help. Eddie's ruse works, and George returns to New York with Eddie to appear in the Ziegfeld Follies. On opening night, George sings "It Had to Be You" to Connie, who is seated in the audience, and after the show, a double wedding is held for Eddie and Joan and George and Connie. As Connie and George leave for their honeymoon in Niagara Falls, Joan carries Eddie over the threshold.